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Although they succeeded, the energy that made Orion Pax into Optimus was gone and hence Optimus was once again his former self. In One Shall Rise, Part 3, Megatron and Optimus Prime ventured to Unicron's spark in order to release the Matrix's energy directly into the chaos bringers spark. During the clash with the latter, Optimus would be forced to contend with the Quintessons' use of a Matrix Replica. Sensing potential and nobility within Orion, Primus gave him the Matrix, thus turning Orion Pax into Optimus Prime. The Matrix would subsequently guide and aid him through a number of adventures, including continued battles with Megatron and the Decepticons and conflicts with new foes such as the Star Seekers and the Quintessons. When Primus was reaching to a critical state, Orion Pax went to Cybertron's core and tried to help Primus. When the High Council named Orion Pax the new Prime and gave him the task of finding the Matrix for himself, Megatron was enraged and vowed to find it for his personal use first. Several had dreams of reclaiming it and becoming the Prime that would lead Cybertron to a new Golden Age. The Matrix became an object of legend among Cyberton. The Matrix was held by Prima, but eventually wound up with Primus in the Core.

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